故事开始于二十世纪20年代的陕南地区。胡家村村民因不满官府苛捐杂税、横征暴敛,与官府发生冲突,遭官兵追杀,全村老少被迫逃到豫西山区狗娃山,自立山寨为伙。大家一致推举大黑骡(范明 饰)为大当家,女当家被尊称为山寨的“奶奶”(宁静 饰)。逃亡途中奶奶捡到一名将要饿死的男孩,为好养活遂起名狗娃(罗晋 饰)。数年后狗娃长大成人,时逢当地遭百年不遇的大旱,张家堡农民因交不起地租遭恶霸吃人贼(李耕 饰)血洗,奶奶率山寨一干人马驰援张家堡。吃人贼之子李冬青(芦芳生 饰)留日归来,继承恶父衣钵,率保安团围剿狗娃山,大当家殒命。狗娃义愤填膺,率众五战五捷,直至抗日战争爆发,和奶奶一起投入到民族解放战争的洪流之中......
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